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The Combine chased the now-Xenians out of their home dimension into Xen, the border world between all dimensions. Their leader, the Nihilanth, was raising an army (the grunts) using his unwilling slaves (the Vortigaunts) to fight the Combine when they came to Xen. Everything alive on Xen came...

I think that when a headcrab mutates its host, what it's doing is assimilating the host's biology to its own. This eventually results in a Gonarch. It doesn't matter what the host is, because it's all turning into headcrab anyway. EDIT: Great guide, by the way. Exactly what people like me need...