VirusType2
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In other words, wheelspin. The amount of friction of a wheel is extremely low, since the contact area is small (and in this case, the wheels freespin to a large degree) Even if the wheels can't keep up, a plane will easily overcome this small amount of friction. However, we aren't talking about frictionless wheels, we are talking about tires, which have good grip - but a treadmill isn't necessarily the ideal surface for a tire, so the plane would very easily overcome this friction.The treadmill will not exert enough force upon the plane's wheels to counteract the acceleration caused by the plane's turbines.
So, if the surface of the tread-mil was equivalent to tarmac, I question whether the wheels would be positioned forward enough to where the plane wouldn't nose into the treadmill. In other words, the balance of the plane would be important, as would the amount of grip on the thread-mil.